“If Planavsky’s estimation of these oxygen levels are correct, it could mean that simple organisms were ready to evolve into complexity for ages before they actually got the chance.”
So in other words, their DNA contained complexities before they evolved. Like fish contained feet DNA years before they grew feet.
It would not make any sense to me how anything could evolve traits in their DNA that would not benefit them immediately.
Or perhaps one wing sprouted 100 million years before the other one.
...or perhaps God is not a Liar...