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Allow me to share a bit. Bill spoke very warmly of his brother-like relationship with Leonard Nimoy, and he was obviously very fond of DeForest Kelley as well - one of his stories was about how he'd watch the mannerisms of people - especially other actors, before they got "into character". Especially Kelley...

One of those observations of Dee Kelley was as he popped a bagel into the toaster at the studio's breakfast kiosk and then stood there radiating eager anticipation. Bill couldn't resist a good prank, though, so he enlisted Nimoy to distract Kelley before the toaster popped up - removing the bagel and pushing the toaster handle back down. Kelley returned in time to see the toaster pop up - empty, and then a range of expressions crossed his face - puzzlement, disbelief, and finally anger when he realized that he'd been Shetner's victim once again.

It's just a tiny peek, but I think that story offers a glimpse into why that particular casting "chemistry" worked.