In nature, the first moment some animal species gets penned into some small area, it most often goes extinct as was the case with the little heath hen I mentioned.
Just because the conditions I alluded to are man-made doesn’t mean other similar conditions don’t occur in nature.
I tend to think mathematically. I can conceive of local fitness maxima that are not global. For example, you might have traits tailored to a particular local environment which tend to evolve at the expense of some others which might more often be useful. “More often useful”is what I think of as globally optimized. Do you look at it differently?