Two New Discoveries Answer Big Questions In Evolution Theory

Wall Street Journal ^ | 07 April 2006 | SHARON BEGLEY

[CarolinaGuitarman]

"The fable of evolution is the religion of the athiest(sic) materialist who desires to throw off all moral laws and indulge in the sexual desires of their arrogant heart."

That's why most people who accept evolution in the USA are Christians.

[AntiGuv]

The fable of evolution is the religion of the athiest materialist who desires to throw off all moral laws and indulge in the sexual desires of their arrogant heart.

Wow. I am definitely not getting my money's worth lately..

[dmz]

Only took 10 posts to get all evolutionists labelled as sex-addicted materialistic atheists doomed to eternal damnation.

Oh, we kick toy poodles and breed pit bull terriers to be vicious as well.

[ConsentofGoverned]

"I don't find anyone referring to the scientists of several hundred years ago as laughing stocks. They did what they could with their tools and accumulated knowledge. Same as today"
>>>>>>>>>>..........'
If I was posting that then you would be right..I was talking about todays dupes who refuse to see that evolution theory does not have the tools/power to say the species diversity was caused by evolution theory alone..you are using the science of 1800's to advance this claim ..todays so called scientist are who will be laughed at.

[protest1]

LOL... how many "missing links" does that make now? Only for them to be quietly forgotten when they turn out to just one more case of wishful thinking like the coelacanth "fish that walked".

[Ichneumon]

One study produced what biblical literalists have been demanding ever since Darwin -- the iconic "missing links." If species evolve, they ask, with one segueing into another, where are the transition fossils, those man-ape or reptile-mammal creatures that evolution posits?

This is worded poorly -- it might give the impression that up until this recent find, such transitional fossils have been lacking. This is incorrect. Vast numbers of transitional fossils in many different lineages have been found over the past 100+ years. For example, this paragraph mentions "reptile-mammal creatures" -- here is a brief overview of the fine-grained transitional sequences which have been found:

Example 2: reptile-mammals

Figure 1.4.1. The jaws of three vertebrates—mammal, therapsid, and pelycosaur. A side view of three idealized skulls of mammals, therapsids (mammal-like reptiles), and pelycosaurs (early reptiles). The figure shows the differences between mammal and reptilian jaws and ear-bone structures. The jaw joint is shown as a large black dot, the quadrate (mammalian anvil or incus) is in turquoise, the articular (mammalian hammer or malleus) is in yellow, and the angular (mammalian tympanic annulus) is in pink. Note how, in the reptile, the jaw joint is formed between the blue quadrate and the yellow articular (with the pink angular close by), and how, in the mammal, the jaw joint is formed between the squamosal above and the dentary below. In the reptile, the squamosal is just above and contacting the quadrate. Advanced therapsids have two jaw joints: a reptile-like joint and a mammal-like joint (Figure based on Kardong 2002, pp. 275, reproduced with permission from the publisher, Copyright © 2002 McGraw-Hill)

We also have an exquisitely complete series of fossils for the reptile-mammal intermediates, ranging from the pelycosauria, therapsida, cynodonta, up to primitive mammalia (Carroll 1988, pp. 392-396; Futuyma 1998, pp. 146-151; Gould 1990; Kardong 2002, pp. 255-275). As mentioned above, the standard phylogenetic tree indicates that mammals gradually evolved from a reptile-like ancestor, and that transitional species must have existed which were morphologically intermediate between reptiles and mammals—even though none are found living today. However, there are significant morphological differences between modern reptiles and modern mammals. Bones, of course, are what fossilize most readily, and that is where we look for transitional species from the past. Osteologically, two major striking differences exist between reptiles and mammals: (1) reptiles have at least four bones in the lower jaw (e.g. the dentary, articular, angular, surangular, and coronoid), while mammals have only one (the dentary), and (2) reptiles have only one middle ear bone (the stapes), while mammals have three (the hammer, anvil, and stapes) (see Figure 1.4.1).

Early in the 20^th century, developmental biologists discovered something that further complicates the picture. In the reptilian fetus, two developing bones from the head eventually form two bones in the reptilian lower jaw, the quadrate and the articular (see the Pelycosaur in Figure 1.4.1). Surprisingly, the corresponding developing bones in the mammalian fetus eventually form the anvil and hammer of the unique mammalian middle ear (also known more formally as the incus and malleus, respectively; see Figure 1.4.2) (Gilbert 1997, pp. 894-896). These facts strongly indicated that the hammer and anvil had evolved from these reptilian jawbones—that is, if common descent was in fact true. This result was so striking, and the required intermediates so outlandish, that many anatomists had extreme trouble imagining how transitional forms bridging these morphologies could have existed while retaining function. Young-earth creationist Duane Gish stated the problem this way:

"All mammals, living or fossil, have a single bone, the dentary, on each side of the lower jaw, and all mammals, living or fossil, have three auditory ossicles or ear bones, the malleus, incus and stapes. ... Every reptile, living or fossil, however, has at least four bones in the lower jaw and only one auditory ossicle, the stapes. ... There are no transitional fossil forms showing, for instance, three or two jawbones, or two ear bones. No one has explained yet, for that matter, how the transitional form would have managed to chew while his jaw was being unhinged and rearticulated, or how he would hear while dragging two of his jaw bones up into his ear." (Gish 1978, p. 80)

Figure 1.4.2. A comparison of the ears of reptiles and mammals. The reptile ear is shown on the left, the mammal ear on the right. As in Figure 1.4.1, the quadrate (mammalian anvil or incus) is in turquoise and the articular (mammalian hammer or malleus) is in yellow. The stapes is shown in brown. Note how the relative arrangement of these bones is similar in both taxa, in the order of inner ear-stapes-quadrate-articular.

Gish was incorrect in stating that there were no transitional fossil forms, and he has been corrected on this gaff numerous times since he wrote these words. However, Gish's statements nicely delineate the morphological conundrum at hand. Let's review the required evolutionary conclusion. During their evolution, two mammalian middle ear bones (the hammer and anvil, aka malleus and incus) were derived from two reptilian jawbones. Thus there was a major evolutionary transition in which several reptilian jawbones (the quadrate, articular, and angular) were extensively reduced and modified gradually to form the modern mammalian middle ear. At the same time, the dentary bone, a part of the reptilian jaw, was expanded to form the major mammalian lower jawbone. During the course of this change, the bones that form the hinge joint of the jaw changed identity. Importantly, the reptilian jaw joint is formed at the intersection of the quadrate and articular whereas the mammalian jaw joint is formed at the intersection of the squamosal and dentary (see Figure 1.4.1).

How could hearing and jaw articulation be preserved during this transition? As clearly shown from the many transitional fossils that have been found (see Figure 1.4.3), the bones that transfer sound in the reptilian and mammalian ear were in contact with each other throughout the evolution of this transition. In reptiles, the stapes contacts the quadrate, which in turn contacts the articular. In mammals, the stapes contacts the incus, which in turn contacts the malleus (see Figure 1.4.2). Since the quadrate evolved into the incus, and the articular evolved into the malleus, these three bones were in constant contact during this impressive evolutionary change. Furthermore, a functional jaw joint was maintained by redundancy—several of the intermediate fossils have both a reptilian jaw joint (from the quadrate and articular) and a mammalian jaw joint (from the dentary and squamosal). Several late cynodonts and Morganucodon clearly have a double-jointed jaw. In this way, the reptilian-style jaw joint was freed to evolve a new specialized function in the middle ear. It is worthy of note that some modern species of snakes have a double-jointed jaw involving different bones, so such a mechanical arrangement is certainly possible and functional.

Since Figure 1.4.3 was made, several important intermediate fossils have been discovered that fit between Morganucodon and the earliest mammals. These new discoveries include a complete skull of Hadrocodium wui (Luo et al. 2001) and cranial and jaw material from Repenomamus and Gobiconodon (Wang et al. 2001). These new fossil finds clarify exactly when and how the malleus, incus, and angular completely detached from the lower jaw and became solely auditory ear ossicles.

Recall that Gish stated: "There are no transitional fossil forms showing, for instance, three or two jawbones, or two ear bones" (Gish 1978, p. 80). Gish simply does not understand how gradual transitions happen (something he should understand, obviously, if he intends to criticize evolutionary theory). These fossil intermediates illustrate why Gish's statement is a gross mischaracterization of how a transitional form should look. In several of the known intermediates, the bones have overlapping functions, and one bone can be called both an ear bone and a jaw bone; these bones serve two functions. Thus, there is no reason to expect transitional forms with intermediate numbers of jaw bones or ear bones. For example, in Morganucodon, the quadrate (anvil) and the articular (hammer) serve as mammalian-style ear bones and reptilian jaw bones simultaneously. In fact, even in modern reptiles the quadrate and articular serve to transmit sound to the stapes and the inner ear (see Figure 1.4.2). The relevant transition, then, is a process where the ear bones, initially located in the lower jaw, become specialized in function by eventually detaching from the lower jaw and moving closer to the inner ear.

Figure 1.4.3. A comparison of the jawbones and ear-bones of several transitional forms in the evolution of mammals. Approximate stratigraphic ranges of the various taxa are indicated at the far left (more recent on top). The left column of jawbones shows the view of the left jawbone from the inside of the mouth. The right column is the view of the right jawbone from the right side (outside of the skull). As in Figure 1.4.1, the quadrate (mammalian anvil or incus) is in turquoise, the articular (mammalian hammer or malleus) is in yellow, and the angular (mammalian tympanic annulus) is in pink. For clarity, the teeth are not shown, and the squamosal upper jawbone is omitted (it replaces the quadrate in the mammalian jaw joint, and forms part of the jaw joint in advanced cynodonts and Morganucodon). Q = quadrate, Ar = articular, An = angular, I = incus (anvil), Ma = malleus (hammer), Ty = tympanic annulus, D = dentary. (Reproduced from Kardong 2002, pp. 274, with permission from the publisher, Copyright © 2002 McGraw-Hill)

The above is from 29+ Evidences for Macroevolution, which compiles several hundred transitional fossils, which is itself just a *SMALL* sampling of the ENORMOUS numbers of fine transitional sequences found in the fossil record and well known to anyone who has bothered to CRACK OPEN A BOOK -- or even do a websearch -- in the past 25 years or so... So what's the anti-evolutionists' excuse for remaining abysmally ignorant of such things, and repeatedly making the false claim that there are "no" transitional fossils, etc.?

Here's another look:

Mammal-Like Reptiles

As previously stated, a succession of transitional fossils exists that link reptiles (Class Reptilia) and mammals (Class Mammalia). These particular reptiles are classifie as Subclass Synapsida. Presently, this is the best example of th e transformation of one major higher taxon into another. The morphologic changes that took place are well documented by fossils, beginning with animals essentially 100% reptilian and resulting in animals essentially 100% mammalian. Therefore, I have chosen this as the example to summarize in more detail (Table 1, Fig. 1).

Skulls and jaws of synapsid reptiles and mammals; left column side view of skull; center column top view of skull; right column side view of lower jaw. Hylonomus modified from Carroll (1964, Figs. 2,6; 1968, Figs. 10-2, 10-5; note that Hylonomus is a protorothyrod, not a synapsid). Archaeothyris modified from Reisz (1972, Fig. 2). Haptodus modified from Currie (1977, Figs, 1a, 1b; 1979, Figs. 5a, 5b). Sphenacodo n modified from Romer & Price (1940, Fig. 4f), Allin (1975, p. 3, Fig. 16);note: Dimetrodon substituted for top view; modified from Romer & Price, 1940, pl. 10. Biarmosuchus modified from Ivakhnenko et al. (1997, pl. 65, Figs. 1a, 1B, 2); Alin & Hopson (1992; Fig. 28.4c); Sigogneau & Tchudinov (1972, Figs. 1, 15). Eoarctops modified from Broom (1932, Fig. 35a); Boonstra (1969, Fig. 18). Pristerognathus modified from Broom (1932, Figs 17a, b,c); Boonstra (1963, Fig. 5d). Procynosuchus modified from Allin & Hopson (1992, Fig. 28.4e); Hopson (1987, Fig. 5c); Brink (1963, Fig. 10a); Kemp (1979, Fig. 1); Allin (1975, p. 3, Fig. 14). Thrinaxodon modified from Allin & Hopson (1992, Fig. 28.4f);Parrington (1946, Fig. 1); Allin (1975, p. 3, Fig. 13). Probainognathus modified from Allin & Hopson (1992, Fig. 28.4g); Romer (1970, Fig. 1); Allin (1975, p. 3, Fig. 12). Morga nucodon modified from Kermack, Mussett, & Rigney (1981, Figs. 95, 99a; 1973, Fig. 7a); Allin (1975, p. 3, Fig. 11). Asioryctes modified from Carroll (1988, Fig. 20-3b). Abbreviations: ag = angular; ar = articular; cp = coronoid process; d = dentary; f = lateral temporal fenestra; j = jugal; mm = attachment site for mammalian jaw muscles; o = eye socket; po = post orbital; q = quadrate; rl = reflected lamina; sq = squamosal; ty = tympanic.

TAXONOMY	LATERAL TEMPORAL FENESTRA	LOWER JAW DENTARY	TEETH	LOWER JAW: POST- DENTARY BONES	MIDDLE EAR & JAW ARTICULATION
M: Early Placental mammals Asioryctes Upper Cretaceous	Merged with eye socket; cheek arch bowed out laterally	100% of jaw length is the den- tary; condylar process in contact with squamosal	Fully differentiated teeth; incisors, canines, premolars; one tooth replacement	No post-dentary bones	3 middle ear bones (stapes, incus, malleus) + tympanic; squamosal-dentary jaw joint
L: "Pantothere" mammals Amphitherium Middle/Upper Jurassic	X	100% of jaw length is the den-  tary; condylar process contacts squamosal	Fully differentiated teeth; incisors, canines, premolars; one tooth replacement	Post-dentary bones migrated to middle ear	Probably 3 middle ear bones (stapes, incus, malleus) + tympanic; squamosal-dentary jaw joint
K: Morganucodontid mammals Morganucodon  Upper Triassic & Lower Jurassic	Merged with eye socket; cheeck arch bowed out laterally	100% of jaw length is the den- tary; condylar process expanded posteriorly to make contact with squamosal	Fully differentiated teeth; incisors, canines, premolars; one tooth replacement	20% of jaw length; reflected lamina decreased to narrow ribbon-like horseshoe	Stapes extends from inner ear capsule to quadrate; quadrate tiny; both quadrate-articular and squamosal-dentary jaw joints
J: Chiniquodontid cynodonts Probainognathus Middle Triassic	Much larger than eye socket; 40- 45% of skull length; expanded posterioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally	95% of jaw length is the dentary; large coronoid process expanded posteriorly; condylar process expanded posteriorly	Large single canine; cheek teeth multicusped; tooth replacement reduced	20% of jaw length; angular notch widened ventrally; width of main part of angular decreased; reflec - ted lamina decreased to narrow ribbon-like horseshoe	Stapes extends from inner ear capsule to quadrate; quadrate tiny; quadrate-articular joint
I:Galesaurid cynodonts Thrinaxodon Lower Triassic	Much larger than eye socket; 40% of skull length; expanded pos- terioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally	85% of jaw length is the dentary; large coronoid process expanded to top of eye socket and pos- teriorly; jaw muscles attached to most of coronoid process	Large single canine; cheek teeth multicusped; tooth replacement reduced	25% of jaw length; angular notch widened ventrally; width of reflec- ted lamina decreased; width of main part of angular decreased	Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint
H: Procynosuchid cynodonts Procynosuchus upper Upper Permian	Much larger than eye socket; 40% of skull length; expanded pos- terioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally	75-80% of jaw length is the den- tary; coronoid process expanded to near top of eye socket and posteriorly; jaw muscles  attached to dorsal part of coronoid process	Large single canine; cheek teeth multicusped	30% of jaw length; angular notch widened ventrally; width of reflected lamina decreased	Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint
G: Early Therocephalians Pristerognathus lower Upper Permian	Larger than eye socket; expanded posteriorly and medially; 30% of skull length	75-80% of jaw length is the den- tary; posterior end of dentary expanded posteriorly and dorsally into narrow blade-like coronoid process; rises to middle of eye socket	Large single canine; other teeth simple cones.	35% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like	Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint
F: Early Gorgonopsians Eoarctops lower Upper Permian	Slightly larger than eye socket; expanded posteriorly and medially (minimal); 20-25% of skull length	65-75% of jaw length is the den- tary; posterior end of dentary slightly expanded posteriorly and dorsally as incipient coronoid process	Large single canine; other teeth simple cones.	40% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint
E: Eotitanosuchians Sphenacodon Lower Permian	Small; slightly smaller than eye socket; slightly expanded posteriorly and medially	65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row	Large single canine; other teeth simple cones.	40% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like	Stapes extends from inner ear capsule to quadrate;  quadrate- articular jaw joint
D: Late sphenacodonts Sphenacodon Upper Pennsylvanian	Small; smaller than eye socket; confined to one side of skull	65% of jaw length is the dentary; posterodorsal edge rises broadly but slightly above the tooth row	Enlarged incipient canines; other teeth simple cones	60% of jaw length; venntral edge of angular notched ("angular notch") offsetting a short pro- tusion (reflected lamina)	Stapes extends from inner ear capsule to quadrate; quadrate large and plate-like; quadrate- articular jaw joint
C: Early spenacodonts Haptodus Upper Pennsylvanian	Tiny; smaller than eye socket; confined to one side of skull	65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row	Undifferentiated; slightly enlarged incipient canines just behind nares	70% of jaw length; ventral edge of angular with shallow indentation	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint
B: Early ophiacodonts Archaothyris upper Middle Pennsylvanian	Tiny; smaller than eye socket; confined to one side of skull	x	Undifferentiated; slightly enlarged incipient canines just behind nares	x	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint
A: Protorothyrids Hylonomus lower Middle Pennsylvanian	Absent	65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row	Undifferentiated; slightly enlarged incipient canines just behind nares	70% of jaw length; ventral edge of angular continuous	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint

Table 1: Morphology of synapsid reptiles and mammals (Note that Hylonomus is a protothyrid, not a synapsid). Data from references cited in text.

Modern reptiles and mammals are very distinctive, easily diagnosable, and do not intergrade. Reptiles are covered by scales, mammals by hair; reptiles are cold-blooded, mammals warm-blooded; reptiles do not suckle their young, mammals have mammary glands; reptiles have sprawling posture, mammals have upright posture. Most of these features are soft part anatomy or physiology that very rarely fossilize (although dinosaur skin impressions are known from Cretaceous sediments, and imprints of mammal hair are known from Eocene bats from Germany; Franzen, 1990). In the fossil record, we must look to skeletal features.

There are many skeletal features which allow us to distinguish the reptiles from the mammals (Carroll, 1988; Table 1, rows A, M). The single most important defining characteristic is the nature of the articulation of the lower jaw to the skull (Simpson, 1959). In reptiles, multiple bones comprise the lower jaw. A small bone at the posterior end of the lower jaw, the articular, articulates with the quadrate bone of the skull (Simpson, 1959; Carroll, 1988). In mammals, one large bone, the dentary, comprises the lower jaw. It articulates with the squamosal bone of the skull (Simpson, 1959; Carroll, 1988).

From comparative anatomy studies, it is certain that most of the bones of the reptiles and mammals are homologous (Crompton & Parker, 1978; Carroll, 1988). Of greatest importance, the middle ear bones of mammals (stapes, incus, malleus, and tympanic) are homologous with several of the skull and jaw bones of reptiles (stapes, quadrate, articular, and angular, respectively; Romer, 1956, p. 33-38, 1970a; Allin, 1975, 1986; Allin & Hopson, 1992; Crompton & Parker, 1978; Hopso n, 1987, 1994; Carroll, 1988). One group of reptiles, the synapsids (Subclass Synapsida), share with the mammals an additional homologous structure: the lateral temporal fenestra, which is an opening in the skull behind the eye socket at the triple junction between the squamosal, jugal , and post orbital bones (Broom, 1932; Frazetta, 1968; Kemp, 1982; Carroll, 1988). A band of bone composed of the jugal and the squamosal is adjacent to the lateral temporal fenestra (Broom, 1932; Kemp, 1982; Carroll, 1988). This is the cheek arch so characteristic of mammal skulls (Broom, 1932; Kemp, 1982; Carroll, 1988). Therefore, synapsids are commonly named the “mammal-like reptiles.”

The presence of diagnosable morphologic differences between reptiles (including the oldest reptiles and the oldest synapsids) and mammals distinguishes them as distinct taxa. This allows us to test evolution by looking for transitional forms between the two. Because many of the bones are homologous, we should find evidence illustrating how these bones were modified over time to become the new bones. Furthermore, these morphologic changes should happen in parallel and in geochronologic succession.

Synapsid reptiles inhabited Pangea from the Middle Pennsylvanian through the Early Jurassic (Kemp, 1982, 1985; Sloan, 1983; Carroll, 1988; Hopson, 1969, 1987, 1994; Hopson & Crompton, 1969; Hotton, et al., 1986; Crompton & Jenkins, 1973; Sidor & Hopson, 1998; Romer & Price, 1940; Broom, 1932; Boonstra, 1963, 1969, 1971; Tchudinov, 1983; Olson, 1944; Tatarinov, 1974; Vyushkov, 1955; Efremov, 1954). From the Early Permian through the Early Triassic, they were the largest and most abundant land animals (Sloan, 1983; Colbert, 1965). Though much less well known to the general public than dinosaurs, one of the “cereal box dinosaurs,” Dimetrodon (the sail-backed reptile), is a synapsid, not a dinosaur (Romer & Price, 1940; Carroll, 1988). The oldest mammals are Late Triassic (Kemp, 1982; Carroll, 1988). Below is a discussion of the geochronologic succession linking synapsids and mammals. The oldest reptiles (named protorothyrids; Carroll, 1964, 1988, p. 192-199) are from the lower Middle Pennsylvanian, and the oldest synapsids (Reisz, 1972) are from the upper Middle Pennsylvanian, both of Nova Scotia. Upper Pennsylvanian and Lower Permian forms are known primarily from the midcontinent and Permian Basin region of the United States (Romer & Price, 1940; Currie, 1977, 1979; Kemp, 1982; Sloan, 1983). The basal Upper Permian forms are known from Russia (Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Ivakhnenko et al., 1997). Most of the Upper Permian and Lower Triassic succession is known from southern Africa, especially the Great Karoo of South Africa (Broom, 1932; Boonstra, 1963, 1969, 1971; Hopson & Kitching, 1972; Kemp, 1982; Sloan, 1983). The Middle Triassic forms are from South America (Romer, 1969a, 1969b, 1970b, 1973; Romer & Lewis, 1973; Bonaparte & Barbarena, 1975), and the Upper Triassic and Lower Jurassic mammals are known from Eurasia (Kermack, Mussett, & Rigney, 1973, 1981; Kemp, 1982). Subsequent Mesozoic mammals are known from all over the world (Simpson, 1928; Lillegraven et al., 1979).

When placed in proper geochronologic succession, the synapsids naturally form a succession of taxa (genera and families) that progressively become more mammal-like and less reptile-like (Kemp, 1982, 1985; Sloan, 1983; Sidor & Hopson, 1998; Hopson, 1987, 1994). Morphologic changes, summarized in Table 1 and Figure 1, affect the entire skeletal anatomy of these animals, but are most clearly displayed in their skulls.

The lateral temporal fenestra increased in size from a tiny opening smaller than the eye socket to a giant opening occupying nearly half the length of the skull. Ultimately, it merged with the eye socket, thus producing the full development of the cheek arch so characteristic of mammals (Broom, 1932; Frazetta, 1968; Kemp, 1982; Sloan, 1983; Hopson, 1987, 1994; Carroll, 1988).

Successively, the relative proportion of the lower jaw comprised of the dentary bone (teeth-bearing bone) gradually increased until the entire lower jaw consisted of the dentary (Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1987, 1994). In Pennsylvanian and Lower and basal Upper Permian synapsids, the postero-dorsal edge of the lower jaw rose broadly but only slightly above the level of the tooth row (Romer & Price, 1940; Currie, 1977, 1979; Ivakhnenko et al., 1997; Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Hopson, 1987, 1994). In succeeding forms, the posterior part of the dentary expanded dorsally and posteriorly as a blade-like process, and progressively became larger (Broom, 1932; Boonstra, 1963, 1969, 1971; Sigogneau, 1970; Brink, 1963; Kemp, 1979; Hopson, 1987, 1994), forming the coronoid process (Parrington, 1946; Fourie, 1974; Romer, 1969b, 1970b, 1973; Hopson, 1987, 1994) to which the mammalian-type jaw musculature is attached (Barghusen, 1968; Bramble, 1978; Crompton, 1972; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). Concomitantly, the post-dentary bones progressively reduced in size (Allin, 1975; Crompton, 1972; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1987, 1994).

Beginning with the Upper Pennsylvanian sphenacodonts, a notch developed in the angular bone that offsets a projection, the reflected lamina (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Romer & Price, 1940; Currie, 1977, 1979; Kemp, 1982; Sloan, 1983; Carroll, 1988). The reflected lamina first became a large blade-like flange (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Ivakhnenko et al., 1997; Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Broom, 1932; Sigogneau, 1970; Boonstra, 1963, 1969, 1971), and then was progressively reduced to a delicate horseshoe-shaped bone (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Brink, 1963; Parrington, 1946; Fourie, 1974; Romer, 1969b, 1970b, 1973; Kermack, Mussett, & Rigney, 1973, 1981; Kemp, 1979, 1982; Sloan, 1983; Carroll, 1988).

Simultaneously, the quadrate progressively decreased in size (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988). The articular did not decrease in size much, being small initially, but developed a downward-pointing prong (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988). In the synapsids, the lower jaw was hinged to the skull by the articular and quadrate bones (Crompton, 1972; Crompton & Parker, 1978; Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994). Thus they are classified as reptiles (Simpson, 1959; Kemp, 1982; Sloan, 1983; Carroll, 1988). As the quadrate and articular became smaller, they were relieved of their solid suture to the dentary and skull (Crompton, 1972; Allin, 1975, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). A projection of the dentary extended posteriorly and made contact with the squamosal. Morganucodon possessed the mammalian dentary-squamosal jaw joint adjacent to the reptilian articular-quadrate jaw joint (Kermack, Mussett, & Rigney, 1973, 1981; Carroll, 1988). It is classified as the first mammal, but it is a perfect intermediate. Now that a new jaw joint was established, the quadrate and articular were subsequently relieved of that function (Crompton, 1972; Allin, 1975, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). Ultimately, in Middle and Upper Jurassic mammals, the tiny quadrate, articular, and ring-like angular migrated as a unit to the middle ear where they joined the stapes and became the incus, malleus, and tympanic bones (Allin, 197 5, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988).

Progressively, the teeth became differentiated. The large canines developed first, followed by the development of multicusped cheek teeth, reduced tooth replacement (Osborn & Crompton, 1973; Crompton & Parker, 1978), and finally full y differentiated incisors, canines, premolars, and molars with one tooth replacement during life (Kemp, 1982; Hopson, 1994).

Many other morphologic changes are documented in the fossil record. These demonstrate the morphologic and geochronologic succession from sprawling limb posture to upright limb posture of mammals (Jenkins, 1971; Romer & Lewis, 197 3; Kemp, 1982; Carroll, 1988; Hopson, 1994). As Jenkins (1971, p. 210) stated, “In details of morphology and function, the cynodont post-cranial skeleton should be regarded as neither ‘reptilian’ nor ‘mammalian’ but as transitional between the two classes .” Other changes have been adequately summarized elsewhere (Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1994). Obviously, fundamental physiologic changes must have taken place as well, many of which are not directly preserved in the fossil record, though some can be inferred from the skeletal anatomy (Findlay, 1968; Kemp, 1982; Sloan, 1983, Carroll, 1988; Hopson, 1994).

This is well documented in the fossil record by a massive volume of incontrovertible data that cannot be explained away. Such large-scale, progressive, continuous, gradual, and geochronologically successive morphologic change (Sidor & Hopson, 1998) is descent with modification, and provides compelling evidence for evolution on a grand scale.

(The above is from The Fossil Record: Evolution or "Scientific Creation", which is yet ANOTHER source the anti-evolutionists are obviously completely ignorant of -- not that that stops them from spouting off falsehoods about the subject anyway...)

The anti-evolutionists crow about "missing links", and taunt biologists about them, while the biologists just calmly predict, based on the known common ancestry of living creatures, that such intermediates did at one time exist, and will be found in good time. Then when the intermediates *are* found, containing the exact combination of features predicted by evolutionary theory, which provides strong confirmation of that evolutionary theory, the anti-evolutionists either shut up and start complaining about something else, or they wave their hands and cry, "that don't prove nothin'" -- as we've seen already on this thread and the other threads about this recent find and other transitional fossils...

They never get tired of such an intellectually dishonest game.

[Ichneumon]

If I was posting that then you would be right..I was talking about todays dupes who refuse to see that evolution theory does not have the tools/power to say the species diversity was caused by evolution theory alone..you are using the science of 1800's to advance this claim ..todays so called scientist are who will be laughed at.

Yes, your braying laughter based on your gross ignorance of the vast amount of evidence supporting evolutionary biology is a well-known fixture on these threads. Now tell us some more uneducated falsehoods, we get a real kick out of them. Anti-evolutionists make the best Court Jesters.

[Verax]

a previously unknown fish that was on its way to evolving into a four-limbed land-dweller. It had a jaw, fins and scales like a fish, but a skull, neck, ribs and pectoral fin like the earliest limbed animals, called tetrapods.

I plead ignorance here...

Do Darwinian evolutionists theorize that this creature was the very first instance of its species to have the land-dwelling attributes (his mom & dad had no ribs, neck or pectoral fin)...

Or did mom & dad have a half a neck or some other portion of a neck? Or a neck that couldn't hold it's head up? What were the physical manifestation of these transitional anatomies?

How do macroevolutionists like yourselves explain this anotomical progression? What made these transitional lifeforms more fit than their less evolved counterparts?

Thanks.

[ConsentofGoverned]

yada yada yada..the issue is not life has similar characteristics the issue is how did we get the diversity..too many today vote for the random selection of
of chance as the main force with yes millions of years to work..this is pure BS..Life is similar as it was designed that way..simple and true. New species exist because they were created that way from pre existing genetic stock built on pre existing genetic stock..but over in millions of years..the basic question is where did this genetic information come from??answer that and we see ID.

[dmz]

You've missed my point. Several hundred years from now, scientists will look back (as they do now) and recognize that the scientists of today did the best they could with the knowledge and tools available.

If there was another scientific theory available for research purposes on species diversity, it would be done. Are you proposing one? What is it called? What empirical research is being done to further said theory?

You are very strong at knocking down evolution, but I fail to see you proposing your own positive scientific theory.

I don't recall you trumpeting ID as the theory, nor do I recall you quoting Bible verses as though they were scientific evidence, so I am at a loss to determine what you believe accounts for all the biological diversity in evidence on this planet.

[Ichneumon]

LOL... how many "missing links" does that make now?

Several thousand, why do you ask?

Also:

Index to Creationist Claims: Claim CC200: There are no transitional fossils.

Taxonomy, Transitional Forms, and the Fossil Record

On Creation Science and "Transitional Fossils"

The Fossil Record: Evolution or "Scientific Creation"

No transitional fossils? Here's a challenge...

Phylum Level Evolution

Paleontology: The Fossil Record of Life

Cuffey: Transitional Fossils

What Is A Transitional Fossil?

More Evidence for Transitional Fossils

The Origin of Whales and the Power of Independent Evidence

Transitional Forms of Whales

Fossil Horses FAQs

PALAEOS: The Trace of Life on Earth

Mammaliformes: Docodonta

Transitional Fossil Species And Modes of Speciation

Evolution and the Fossil Record

Smooth Change in the Fossil Record

Transitional fossil sequence from dinosaur to bird

Transitional fossil sequence from fish to elephant

Transitional Vertebrate Fossils FAQ

More are being found all the time. For one example, not long ago there were no major transitional fossils between whales and their land-based ancestors. In the time since, however, *many* have been found, mapping out an unmistakable sequence transitioning between land mammals and fully aquatic whales, including this fine fellow:

For details, see:

The Transitional Vertebrate Fossils FAQ

The Origin of Whales and the Power of Independent Evidence

Links on whale evolution

SINE Evolution, Missing Data, and the Origin of Whales

Phylogenetic relationships among cetartiodactyls based on insertions of short and long interpersed elements: Hippopotamuses are the closest extant relatives of whales

Evidence from Milk Casein Genes that Cetaceans are Close Relatives of Hippopotamid Artiodactyls

Analyses of mitochondrial genomes strongly support a hippopotamus±whale clade

A new, diminutive Eocene whale from Kachchh (Gujarat, India) and its implications for locomotor evolution of cetaceans

A new Eocene archaeocete (Mammalia, Cetacea) from India and the time of origin of whales

Mysticete (Baleen Whale) Relationships Based upon the Sequence of the Common Cetacean DNA Satellite1

The Mitochondrial Genome of the Sperm Whale and a New Molecular Reference for Estimating Eutherian Divergence Dates

Limbs in whales and limblessness in other vertebrates: mechanisms of evolutionary and developmental transformation and loss

Eocene evolution of whale hearing

Novel Phylogeny of Whales Revisited but Not Revised

Land-to-sea transition in early whales: evolution of Eocene Archaeoceti (Cetacea) in relation to skeletal proportions and locomotion of living semiaquatic mammals

Subordinal artiodactyl relationships in the light of phylogenetic analysis of 12 mitochondrial protein-coding genes

New Morphological Evidence for the Phylogeny of Artiodactyla, Cetacea, and Mesonychidae

Cetacean Systematics

LIKELIHOOD ESTIMATION OF THE TIME OF ORIGIN OF CETACEA AND THE TIME OF DIVERGENCE OF CETACEA AND ARTIODACTYLA

Phylogenetic Relationships of Artiodactyls and Cetaceans as Deduced from the Comparison of Cytochrome b and 12s rRNA Mitochondrial Sequences

Molecular evolution of mammalian ribonucleases

How many more would you like?

Only for them to be quietly forgotten

They haven't been forgotten at all, except by the anti-evolutionists who like to "forget" how often they've been shown as fools by the discovery of yet another "missing link" they used to bray about, which was then subsequently found.

when they turn out to just one more case of wishful thinking like the coelacanth "fish that walked".

There's no "wishful thinking" about the coelacanth. Are you sure you know what in the hell you're talking about?

[abc1]

There is no proof of anything.
Evolution is just another concoction adhered to by those who want to BELIEVE that we evolved from apes, lizards or some such bottom feeder.
If it were proven we evolved from algae, then what ?
People believe what they want to believe.

[CarolinaGuitarman]

"the issue is not life has similar characteristics the issue is how did we get the diversity..too many today vote for the random selection of
of chance as the main force with yes millions of years to work."

Selection isn't random, by definition.

[From many - one.]

Splurt.

What 1800's genome analysis are you referring to?

These days phenology (appearance) just gives us a direction to start with. Then it's down to the molecular level.

This morning I was reading an analysis of the evolutionary relationships among the sphagnaceae. No phenology involved.

[ConsentofGoverned]

On this subject I did just that in prior threads: proposed ID as a theory that could be verified by standards in current use..just too much to go thru here..you can find the posts on prior ID threads on FR..by the way I accept evolution theory to explain species diversity,,dogs for an example but dogs also show ID..by humans to breed traits we want. that is the basis of my ID theory by the way, another example is modern drug production such as Human insulin from e coli..ID by humans..many here still post ID is not possible LOL

[Verax]

And transitional anamolies with partially functioning anatomies have always been fitter than their peers.

[Verax]

transitional animals

[lemura]

they will be known as a laughing stock as modern methods of gene research prove how ignorant they are on how the multitude of species came into being

While scientists may hold on to pet theories, experience has shown that if a new theory based on either new or existing evidence is developed and gains widespread acceptance, there will be a mad rush to embrace this new theory. Plate tectonics is a very good example.

Absent in the scientific process, however, is faith. It's still all based on evidence, regardless of which theory wins out in the end. The objective is truth, not desire.

[Ichneumon]

yada yada yada..

Keeping to your usual level of discourse, I see.

the issue is not life has similar characteristics

Nor is evolutionary biology based on any evidence as vague and vapid as "life has similar characteristics", if that's that you're trying to imply. If you really think that, you're even more out of your depth in these discussions than I had previously thought.

the issue is how did we get the diversity..

Through evolutionary processes. The evidence for this conclusion is vast and overwhelming, and remains so even though you're completely ignorant of it.

too many today vote for the random selection of of chance as the main force with yes millions of years to work..

...because that's what vast amounts of evidence and research has determined.

this is pure BS..

...you say, based on your deep understanding of biology and your thorough knowledge of the evidence, right? LOL!

Life is similar as it was designed that way..

Feel free to present your evidence for this proposition. If you can manage to produce any, you'll be the first IDer in history to be able to do so. The other have all come up empty-handed so far.

simple and true.

Simple things for simple minds, but there's no evidence indicating that it's "true".

New species exist because they were created that way from pre existing genetic stock built on pre existing genetic stock..but over in millions of years..the basic question is where did this genetic information come from?? P> From evolutionary processes:

Genetic Algorithms

The Origins of Order: Self Organization and Selection in Evolution. By Stuart Kauffman, S. A. (1993) Oxford University Press, NY, ISBN: 0195079515.

Compositional genomes: Prebiotic information transfer in mutually catalytic noncovalent assemblies

Eigen M, and Schuster P, The hypercycle. A principle of natural self-organization. Springer-Verlag, isbn 3-540-09293, 1979

The origin of genetic information: viruses as models

Compositional genomes: prebiotic information transfer in mutually catalytic noncovalent assemblies

Stadler PF, Dynamics of autocatalytic reaction networks. IV: Inhomogeneous replicator networks. Biosystems, 26: 1-19, 1991

Lee DH, Severin K, and Ghadri MR. Autocatalytic networks: the transition from molecular self-replication to molecular ecosystems. Curr Opinion Chem Biol, 1, 491-496, 1997

Lee DH, Severin K, Yokobayashi Y, and Ghadiri MR, Emergence of symbiosis in peptide self-replication through a hypercyclic network. Nature, 390: 591-4, 1997

Apolipoprotein AI Mutations and Information

Creationist Claim CB102: Mutations are random noise; they do not add information.

Multiple Duplications of Yeast Hexose Transport Genes in Response to Selection in a Glucose-Limited Environment

Evolution of biological information

Evolution of biological complexity

Evolution and Information: The Nylon Bug

Examples of Beneficial Mutations and Natural Selection

The evolution of trichromatic color vision by opsin gene duplication in New World and Old World primates

Gene duplications in evolution of archaeal family B DNA polymerases

Koch, AL: Evolution of antibiotic resistance gene function. Microbiol Rev 1981, 45:355378.

Selection in the evolution of gene duplications

Velkov, VV: Gene amplification in prokaryotic and eukaryotic systems. Genetika 1982, 18:529543.

Romero, D & Palacios, R: Gene amplification and genomic plasticity in prokaryotes. Annu Rev Genet 1997, 31:91111.

Stark, GR & Wahl, GM: Gene amplification. Annu Rev Biochem 1984, 53:447491.

Reinbothe, S, Ortel, B, & Parthier, B: Overproduction by gene amplification of the multifunctional arom protein confers glyphosate tolerance to a plastid-free mutant of Euglena gracilis. Mol Gen Genet 1993, 239:416424.

Gottesman, MM, Hrycyna, CA, Schoenlein, PV, Germann, UA, & Pastan, I: Genetic analysis of the multidrug transporter. Annu Rev Genet 1995, 29:607649.

Schwab, M: Oncogene amplification in solid tumors. Semin Cancer Biol 1999, 9:319325.

Widholm, JM, Chinnala, AR, Ryu, JH, Song, HS, Eggett, T, & Brotherton, JE: Glyphosate selection of gene amplification in suspension cultures of three plant species. Physiol Plant 2001, 112:540545.

Otto, E, Young, JE, & Maroni, G: Structure and expression of a tandem duplication of the Drosophila metallothionein gene. Proc Natl Acad Sci USA 1986, 83:60256029.

Maroni, G, Wise, J, Young, JE, & Otto, E: Metallothionein gene duplications and metal tolerance in natural populations of Drosophila melanogaster. Genetics 1987, 117:739744.

Kondratyeva, TF, Muntyan, LN, & Karvaiko, GI: Zinc-resistant and arsenic-resistant strains of Thiobacillus ferrooxidans have increased copy numbers of chromosomal resistance genes. Microbiology 1995, 141:11571162.

Tohoyama, H, Shiraishi, E, Amano, S, Inouhe, M, Joho, M, & Murayama, T: Amplification of a gene for metallothionein by tandem repeat in a strain of cadmium-resistant yeast cells. FEMS Microbiol Lett 1996, 136:269273.

Sonti, RV & Roth, JR: Role of gene duplications in the adaptation of Salmonella typhimurium to growth on limiting carbon sources. Genetics 1989, 123:1928.

Brown, CJ, Todd, KM, & Rosenzweig, RF: Multiple duplications of yeast hexose transport genes in response to selection in a glucose-limited environment. Mol Biol Evol 1998, 15:931942.

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answer that and we see ID.

It's already answered, and no "ID" is seen. Nice try.

[CarolinaGuitarman]

"There is no proof of anything."

Proof is for math and whiskey. That doesn't mean we can't be highly confident in out knowledge of how the physical world works.

"Evolution is just another concoction adhered to by those who want to BELIEVE that we evolved from apes, lizards or some such bottom feeder."

Actually, it's a highly verified ( on a wide number of fronts) scientific theory, accepted by people who look at the data and make logical conclusions. There is no *desire* to have been evolved; it's a fact of life to be accepted.

"If it were proven we evolved from algae, then what ?"

We would know more about the world we live in.

"People believe what they want to believe."

Unfortunately true in far too many cases. That is why science education is so important; it teaches people to not believe what they want to believe but what the evidence dictates. Reality doesn't care what we wish it to be.